| 1 | Harvard University | United States of America (USA) | 173.80 | University of Zurich (UZH) | Switzerland | 21.14 | 194.94 |
| 2 | Chinese Academy of Sciences (CAS) | China | 147.80 | Max Planck Society | Germany | 32.85 | 180.66 |
| 3 | Harvard University | United States of America (USA) | 170.02 | Free University of Berlin (FU Berlin) | Germany | 7.71 | 177.73 |
| 4 | Harvard University | United States of America (USA) | 167.08 | Humboldt University of Berlin (HU Berlin) | Germany | 7.17 | 174.25 |
| 5 | Radboud University Nijmegen (RU) | Netherlands | 88.46 | Max Planck Society | Germany | 82.59 | 171.05 |
| 6 | Harvard University | United States of America (USA) | 85.54 | University of Toronto (U of T) | Canada | 64.88 | 150.42 |
| 7 | Harvard University | United States of America (USA) | 80.66 | Max Planck Society | Germany | 64.54 | 145.20 |
| 8 | National University of Singapore (NUS) | Singapore | 90.21 | Duke University | United States of America (USA) | 53.93 | 144.14 |
| 9 | UCL | United Kingdom (UK) | 85.29 | Max Planck Society | Germany | 54.55 | 139.84 |
| 10 | Max Planck Society | Germany | 96.17 | French National Centre for Scientific Research (CNRS) | France | 36.27 | 132.44 |
| 11 | Harvard University | United States of America (USA) | 88.91 | French National Centre for Scientific Research (CNRS) | France | 27.44 | 116.35 |
| 12 | Harvard University | United States of America (USA) | 67.38 | University of Oxford | United Kingdom (UK) | 48.12 | 115.51 |
| 13 | Harvard University | United States of America (USA) | 79.90 | Helmholtz Association of German Research Centres | Germany | 34.81 | 114.71 |
| 14 | Harvard University | United States of America (USA) | 63.22 | University of Cambridge | United Kingdom (UK) | 47.04 | 110.26 |
| 15 | University of Oxford | United Kingdom (UK) | 61.25 | Max Planck Society | Germany | 44.46 | 105.71 |
| 16 | Harvard University | United States of America (USA) | 59.38 | Chinese Academy of Sciences (CAS) | China | 43.80 | 103.18 |
| 17 | Harvard University | United States of America (USA) | 77.47 | National Institute for Health and Medical Research (INSERM) | France | 20.27 | 97.75 |
| 18 | Harvard University | United States of America (USA) | 55.81 | Karolinska Institute (KI) | Sweden | 39.48 | 95.29 |
| 19 | National Institutes of Health (NIH) | United States of America (USA) | 47.12 | University of Oxford | United Kingdom (UK) | 42.18 | 89.30 |
| 20 | Wellcome Sanger Institute | United Kingdom (UK) | 57.20 | European Molecular Biology Laboratory (EMBL) | Germany | 28.87 | 86.07 |
| 21 | University of Cambridge | United Kingdom (UK) | 57.22 | European Molecular Biology Laboratory (EMBL) | Germany | 25.52 | 82.74 |
| 22 | University of Oxford | United Kingdom (UK) | 64.24 | French National Centre for Scientific Research (CNRS) | France | 16.44 | 80.68 |
| 23 | Helmholtz Association of German Research Centres | Germany | 48.06 | French National Centre for Scientific Research (CNRS) | France | 26.15 | 74.21 |
| 24 | National Institutes of Health (NIH) | United States of America (USA) | 37.97 | University of Cambridge | United Kingdom (UK) | 35.05 | 73.01 |
| 25 | Harvard University | United States of America (USA) | 46.30 | UCL | United Kingdom (UK) | 26.31 | 72.61 |
